March 1989 B.S. (Biology) Kanazawa University, Japan
March 1991 M.S. (Zoology) Kyoto University, Japan
March 1994 D.Sc. (Zoology) Kyoto University, Japan

January 1992-March 1994 Predoctoral Fellow (the Japan Society for the Promotion of Science)
Department of Zoology, Faculty of Science, Kyoto University, Japan

April 1994-September 1995 Postdoctoral Researcher
Bodega Marine Laboratory, University of California at Davis, California, USA

October 1995-June 2000 Instructor
Division of Biological Sciences, Graduate School of Science, Hokkaido University, Japan

July 2000-March 2002 Associate Professor
Department of Life Science, Faculty of Science, Himeji Institute of Technology, Japan

April 2002-March 2004 Associate Professor
Department of Life Science, Graduate School of Science, Himeji Institute of Technology, Japan

April 2004-March 2009 Associate Professor
Department of Life Science, Graduate School of Life Science, University of Hyogo, Japan

April 2009-present Professor
Department of Biology, Faculty of Science and Engineering, Konan University, Japan